Blog List

Wednesday, 25 October 2017

Softwood Anatomy

Author
by Eric Meier ©

Amidst the daunting jungle of hardwood species, (where there are several hundred commercial species used worldwide, with many thousands more that are intermittently seen in woodworking applications), the world of conifers is much more limited. With perhaps only a few dozen chief species worldwide, and only a few hundred species (rather than thousands) that are ever harvested for wood, softwoods represent a comparatively smaller percentage of the diversity seen in wood.
However, softwoods are also much simpler than hardwoods in their anatomical features, so there are far fewer clues to help identify them. Consequently, macroscopic softwood identification is usually less conclusive, and many times will only result in more broad or generic identifications, such as, pine (Pinus spp.) or spruce (Picea spp.), rather than determining an exact species.
The primary distinguishing feature between softwoods and hardwoods is that softwood species lack pores. So upon initial examination of an unknown wood sample, the most immediate detail to note is whether or not the endgrain section has numerous circular openings—commonly referred to as vessels or pores.
With hardwoods, the most conspicuous anatomical feature is the pores, which serves to initially separate hardwoods based upon the various distributions of their pores. However, because softwoods are poreless, another conspicuous and easily recognizable anatomical feature is used instead to quickly make broad distinctions between conifers.

Resin Canals

Resin canals, sometimes referred to as resin ducts, are unique to conifers; they are technically not individual cells, but are actually open, tube-like spaces bordered by special cells that have the ability to secrete pitch or resin into the neighboring opening (canal). One of the apparent purposes of these ducts is to protect and seal up a wound by exuding resin to cover the damaged area of the tree. (Accordingly, in rare instances, some softwood species that don’t normally have resin ducts will develop them as a result of trauma.)
When initially sorting between softwoods, three primary groups emerge when considering their resin canals:
Large, numerous, evenly spaced, solitary resin canalsSmall, infrequent, sporadically spaced, tangentially-grouped resin canalsNo resin canals

Ponderosa Pine (endgrain 10x)

Sitka Spruce (endgrain 10x)

Cedar of Lebanon (endgrain 10x)
This group includes all of the true pines in the Pinus genus.Some genera in the Pinaceae family, such as: spruce (Picea spp.), larch (Larix spp.), and Douglas-Fir (Pseudotsuga spp.)Numerous species of cedar, as well as: fir (Abies spp.), hemlock (Tsuga spp.), juniper (Juniperus spp.), and yew (Taxus spp.)

Resin canals can be very infrequent in some softwood species, so care must be taken to observe an adequately large area of the endgrain to ensure their absence. One way to confirm the presence of resin canals is to look for a type of horizontal resin canal, called a fusiform ray. When viewed from the endgrain, fusiform rays are seen as unusually wide rays, and can be thought of as a hybrid between a resin canal and a ray. (See the picture of Sitka Spruce above for an example of fusiform rays.)
Once a softwood sample has been sorted into one of the three initial categories based upon the resin canals, further refinements are used to obtain a more specific and positive identification. The following is an overview of the remaining features that are used to describe and identify conifers; each softwood species’ profile page also lists these characteristics to help in identification.

Tracheids

Within a conifer’s trunk, the majority of the wood is comprised of long, thin cells called tracheids. In addition to giving the tree most of its strength, tracheids also double as “pores” in a sense—since conifers lack true pores, they rely on tracheids for sap conduction.
Tracheids are arranged in orderly, neat rows within the trunk, and are longitudinal (they run lengthwise with the trunk). Tracheids can vary in diameter depending on the species, and because they account for over 90% of the cells, their diameter greatly influences the overall texture and feel of the wood—the larger the diameter, the coarser the texture.

Tracheid Diameter and Texture

Relative SizeMicrons (µm)Texture
Very Small< 25Very Fine
Small25-35Fine
Medium35-50Medium
Large50-60Coarse
Very Large> 60Very Coarse

Because anatomical features such as tracheids are very difficult to accurately measure without specialized tools, their diameter will instead be referred to in comparative terms, such as small, medium, and large. When viewing the endgrain under 10x magnification, softwood species with the largest tracheids generally have cells that can each be seen individually. Single cells become harder to discern in the mid-range, while the smallest tracheids tend to appear only as a collective mass with no clear cell borders.
Cypress (endgrain 10x)

Balsam Fir (endgrain 10x)

Northern White Cedar (endgrain 10x)
[Click each picture to get an enlargement with a clearer view of the tracheids] At left is Cypress (Taxodium distichum) with very large tracheids, especially among its earlywood. In the center is Balsam Fir (Abies balsamea) with an intermediate texture. At right is Northern White Cedar (Thuja occidentalis) with very small diameter tracheids that generally can’t be discerned individually.

Earlywood to Latewood Transition

The earlywood of a tree consists of faster-growing material, while the latewood is comprised of denser, slower-growing material. In softwoods, this translates into larger diameter tracheids in the earlywood, and smaller diameter tracheids in the latewood.
In addition to simply noting the overall average size of the tracheids, it can also be helpful to note the rate of transition from the earlywood to the latewood. This transition can be described as being either very gradual, or very abrupt, or somewhere in between, such as moderately abrupt. Although this visual description is somewhat subjective, such observations can help give clues and point an otherwise ambiguous identification into one direction or another.
Southern Yellow Pine (endgrain 10x)
Sugar Pine (endgrain 10x)
Sugar Pine (endgrain 10x)
On the left is Southern Yellow Pine (Pinus spp.), which has a rather abrupt transition from earlywood to latewood. On the right is Sugar Pine (Pinus lambertiana), which has a much more gradual transition in its tracheid sizes.
One caveat in examining the transition rate of the tracheids is that growth rates should also be considered. For instance, if a wood sample has an abnormally fast growth rate, the earlywood and latewood zones will be elongated, giving an artificially lower impression of there being a more gradual transition. Likewise, if a wood sample is encountered that is suspected to have an unusually slow growth rate, then the growth zones will be compressed, and the transition between earlywood and latewood will seem more abrupt.

Grain Contrast

In conifers, grain contrast is basically a description of the contrast of color between the earlywood and latewood. If the color of the wood is somewhat monotone and uninterrupted between growth ring zones, it has low grain contrast. If the color of the wood has a striped appearance with high contrast between the earlywood and latewood, it has high grain contrast.
Much like earlywood to latewood transitions, grain contrast is a somewhat subjective measure—though grain contrast tends to be an easier characteristic to gauge, and can be observed from the face grain as well as the endgrain. Grain contrast can be described as being very high, very low, or anywhere in between.
Western Larch (endgrain 10x)
Western Larch (endgrain 10x)
Eastern White Pine (endgrain 10x)
Eastern White Pine (endgrain 10x)
On the left is a sample of Western Larch (Larix occidentalis) that has high grain contrast. On the right is Eastern White Pine (Pinus strobus), which has a much lower grain contrast; on the facegrain surface of the wood, Eastern White Pine has a very uniform and monotone appearance, with grain lines that can hardly be seen.

It should be noted that grain contrast and earlywood to latewood transitions ought to be considered each separately. Although it would seem apparent that wood with an abrupt transition from earlywood to latewood would also have an uneven grain appearance, it’s important to remember that just because the change in tracheid diameter is abrupt doesn’t necessarily mean that there will be an accompanying change in color—some woods have abrupt transitions, and relatively even grain colors.

Parenchyma

Parenchyma cells are an example of an anatomical element that is of much greater use in macroscopic hardwood identification than it is in softwood identification. Nonetheless, even though hardwood parenchyma cells can form much more complex (and telling) arrangements, softwood parenchyma still have a limited use in identification.
Much like tracheids, parenchyma are oriented along the length of the tree-trunk, and are sometimes referred to as longitudinal or axial parenchyma. In softwoods, these parenchyma cells are roughly the same diameter as tracheids when viewed from the endgrain.
Because tracheids account for over 90% of a softwood’s cells, parenchyma cells are comparatively sparse—(many softwoods don’t have any parenchyma at all, such as: pine, spruce, and yew).
Commonly, single cells that are scattered throughout the cell structure of the wood can’t be individually discerned, but since parenchyma are often filled with darker-colored contents (usually amber or reddish-brown), these special cells can sometimes be spotted with just 10x magnification.
In macroscopic examination of a conifer’s parenchyma, two different formations are typically encountered. The first is when parenchyma is simply scattered throughout the cell structure of the wood in little to no apparent order, which is called diffuse parenchyma. The second is when the cells are arranged in a ring or line that runs parallel to the growth rings, which is frequently called zonate parenchyma.
Redwood (endgrain 10x)
Redwood (endgrain 10x)
Aromatic Red Cedar (endgrain 10x)
Aromatic Red Cedar (endgrain 10x)
On the left is Redwood (Sequoia sempervirens), which has numerous parenchyma cells scattered throughout the wood. On the right, Eastern Red Cedar (Juniperus virginiana) has zonate parenchyma which forms broken lines of reddish cells between the growth ring boundaries. Also note a peculiarity found in this species: the zonate parenchyma also has a tendency to form double-rings between some growth rings.

Rays

With the exception of special fusiform rays, which occur in conjunction with resin canals, regular softwood rays are usually only one to two cells wide (called uniseriate and biseriate, respectively). Because these normal rays are much narrower, and lack unique characteristics that are observable with a 10x hand lens, their usefulness in identification is essentially limited to microscopic examination.
In hardwood identification, rays can offer helpful clues in determining a wood species at only 10x magnification. However, with the scope of this website being limited to macroscopic identification up to only 10x, the minute differences encountered in the narrower rays of the softwood species will not be considered.
For further information log on website :"
http://www.wood-database.com/wood-articles/softwood-anatomy/

No comments:

Post a Comment

Advantages and Disadvantages of Fasting for Runners

Author BY   ANDREA CESPEDES  Food is fuel, especially for serious runners who need a lot of energy. It may seem counterintuiti...